isolated operculum
Thoracosphaera operculata Bramlette & Martini, 1964
The bloom of Cervisiella operculata is recovered from intervals immediately below (latest Maastrichtian) to immediately above (Danian) the Cretaceous–Paleocene mass extinction.
Bramlette, M. N. & Martini, E. 1964. The great change in calcareous nannoplankton fossils between the Maastrichtian and Danian. Micropaleontology. 10(2): 291-322.
Streng, M., Hildebrand-Habel, T. & Willems, H. 2004a. A proposed classification of archeopyle types in calcareous dinoflagellate cysts. Journal of Paleontology. 78(3): 456-483.
Thoracosphaera operculata
Bramlette & Martini, 1964
Danian
McBryde Member, Clayton Formation, Midway Group, near Livingston, Alabama, USA