Set number: 2644

  • Mulleria tuzuniorum 62927 1
  • Mulleria tuzuniorum 62947 2
  • Mulleria tuzuniorum 62965 3
  • Mulleria tuzuniorum 62926 4
  • Mulleria tuzuniorum 62928 5
  • Mulleria tuzuniorum 62946 6
  • Mulleria tuzuniorum 62950 7
  • Mulleria tuzuniorum 62925 8
  • Mulleria tuzuniorum 62953 9
  • Mulleria tuzuniorum 62948 10
  • Mulleria tuzuniorum 62933 11
  • Mulleria tuzuniorum 62967 12
  • Mulleria tuzuniorum 62954 13
  • Mulleria tuzuniorum 62949 14
  • Mulleria tuzuniorum 62931 15
  • Mulleria tuzuniorum 62961 16
  • Mulleria tuzuniorum 62959 17
  • Mulleria tuzuniorum 62943 18
  • Mulleria tuzuniorum 62944 19
  • Mulleria tuzuniorum 62952 20
  • Mulleria tuzuniorum 62957 21
  • Mulleria tuzuniorum 62942 22
  • Mulleria tuzuniorum 62945 23
  • Mulleria tuzuniorum 62951 24
  • Mulleria tuzuniorum 62940 25
  • Mulleria tuzuniorum 62941 26
  • Mulleria tuzuniorum 62939 27
  • Mulleria tuzuniorum 62938 28
  • Mulleria tuzuniorum 62935 29
  • Mulleria tuzuniorum 62936 30
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Holotype in the mobile mount.

Set number: 2651

  • Mulleria tuzuniorum 52983 1
  • Mulleria tuzuniorum 52976 2
  • Mulleria tuzuniorum 52979 3
  • Mulleria tuzuniorum 52981 4
  • Mulleria tuzuniorum 52984 5
  • Mulleria tuzuniorum 52977 6
  • Mulleria tuzuniorum 52978 7
  • Mulleria tuzuniorum 52980 8
  • Mulleria tuzuniorum 52996 9
  • Mulleria tuzuniorum 52986 10
  • Mulleria tuzuniorum 52997 11
  • Mulleria tuzuniorum 52990 12
  • Mulleria tuzuniorum 52995 13
  • Mulleria tuzuniorum 52985 14
  • Mulleria tuzuniorum 52998 15
  • Mulleria tuzuniorum 52989 16
  • Mulleria tuzuniorum 52991 17
  • Mulleria tuzuniorum 52988 18
    10µm
Carlamuelleria tuezueniorum, Middle Paleocene, DSDP Leg 39, Sites 354, Ceará Rise, North Atlantic Ocean

In the mobile mount.

Set number: 2650

  • Mulleria tuzuniorum 65398 1
  • Mulleria tuzuniorum 65410 2
  • Mulleria tuzuniorum 65401 3
  • Mulleria tuzuniorum 65403 4
  • Mulleria tuzuniorum 65399 5
  • Mulleria tuzuniorum 65411 6
  • Mulleria tuzuniorum 65400 7
  • Mulleria tuzuniorum 65405 8
  • Mulleria tuzuniorum 65409 9
  • Mulleria tuzuniorum 65408 10
  • Mulleria tuzuniorum 65404 11
  • Mulleria tuzuniorum 65407 12
  • Mulleria tuzuniorum 65392 13
  • Mulleria tuzuniorum 65384 14
  • Mulleria tuzuniorum 65396 15
  • Mulleria tuzuniorum 65389 16
  • Mulleria tuzuniorum 65391 17
  • Mulleria tuzuniorum 65383 18
  • Mulleria tuzuniorum 65395 19
  • Mulleria tuzuniorum 65390 20
    10µm
Carlamuelleria tuezueniorum, Middle Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount.

Set number: 2649

  • Mulleria tuzuniorum 52922 1
  • Mulleria tuzuniorum 52921 2
  • Mulleria tuzuniorum 52917 3
  • Mulleria tuzuniorum 52918 4
  • Mulleria tuzuniorum 52923 5
  • Mulleria tuzuniorum 52920 6
  • Mulleria tuzuniorum 52916 7
  • Mulleria tuzuniorum 52919 8
  • Mulleria tuzuniorum 52937 9
  • Mulleria tuzuniorum 52933 10
  • Mulleria tuzuniorum 52925 11
  • Mulleria tuzuniorum 52929 12
  • Mulleria tuzuniorum 52936 13
  • Mulleria tuzuniorum 52934 14
  • Mulleria tuzuniorum 52926 15
  • Mulleria tuzuniorum 52930 16
    10µm
Carlamuelleria tuezueniorum, Middle Paleocene, DSDP Leg 39, Sites 354, Ceará Rise, North Atlantic Ocean

In the mobile mount.

Set number: 2648

  • Mulleria tuzuniorum 63125 1
  • Mulleria tuzuniorum 63126 2
  • Mulleria tuzuniorum 63117 3
  • Mulleria tuzuniorum 63120 4
  • Mulleria tuzuniorum 63124 5
  • Mulleria tuzuniorum 63127 6
  • Mulleria tuzuniorum 63119 7
  • Mulleria tuzuniorum 63123 8
  • Mulleria tuzuniorum 63138 9
  • Mulleria tuzuniorum 63140 10
  • Mulleria tuzuniorum 63135 11
  • Mulleria tuzuniorum 63132 12
  • Mulleria tuzuniorum 63136 13
  • Mulleria tuzuniorum 63141 14
  • Mulleria tuzuniorum 63134 15
  • Mulleria tuzuniorum 63130 16
  • Mulleria tuzuniorum 63105 17
  • Mulleria tuzuniorum 63102 18
  • Mulleria tuzuniorum 63147 19
  • Mulleria tuzuniorum 63112 20
  • Mulleria tuzuniorum 63106 21
  • Mulleria tuzuniorum 63104 22
  • Mulleria tuzuniorum 63148 23
  • Mulleria tuzuniorum 63111 24
  • Mulleria tuzuniorum 63109 25
  • Mulleria tuzuniorum 63146 26
  • Mulleria tuzuniorum 63116 27
  • Mulleria tuzuniorum 63108 28
  • Mulleria tuzuniorum 63110 29
  • Mulleria tuzuniorum 63145 30
  • Mulleria tuzuniorum 63115 31
  • Mulleria tuzuniorum 63107 32
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount

Set number: 2647

  • Mulleria tuzuniorum 23743 1
  • Mulleria tuzuniorum 23724 2
  • Mulleria tuzuniorum 23729 3
  • Mulleria tuzuniorum 23728 4
  • Mulleria tuzuniorum 23744 5
  • Mulleria tuzuniorum 23726 6
  • Mulleria tuzuniorum 23727 7
  • Mulleria tuzuniorum 23725 8
  • Mulleria tuzuniorum 23750 9
  • Mulleria tuzuniorum 23747 10
  • Mulleria tuzuniorum 23742 11
  • Mulleria tuzuniorum 23732 12
  • Mulleria tuzuniorum 23749 13
  • Mulleria tuzuniorum 23748 14
  • Mulleria tuzuniorum 23739 15
  • Mulleria tuzuniorum 23733 16
  • Mulleria tuzuniorum 23746 17
  • Mulleria tuzuniorum 23745 18
  • Mulleria tuzuniorum 23735 19
  • Mulleria tuzuniorum 23734 20
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount

Set number: 2646

  • Mulleria tuzuniorum 62088 1
  • Mulleria tuzuniorum 62085 2
  • Mulleria tuzuniorum 62042 3
  • Mulleria tuzuniorum 62041 4
  • Mulleria tuzuniorum 62087 5
  • Mulleria tuzuniorum 62083 6
  • Mulleria tuzuniorum 62043 7
  • Mulleria tuzuniorum 62040 8
  • Mulleria tuzuniorum 62101 9
  • Mulleria tuzuniorum 62100 10
  • Mulleria tuzuniorum 62099 11
  • Mulleria tuzuniorum 62097 12
  • Mulleria tuzuniorum 62096 13
  • Mulleria tuzuniorum 62095 14
  • Mulleria tuzuniorum 62094 15
  • Mulleria tuzuniorum 62093 16
  • Mulleria tuzuniorum 62071 17
  • Mulleria tuzuniorum 62068 18
  • Mulleria tuzuniorum 62067 19
  • Mulleria tuzuniorum 62063 20
  • Mulleria tuzuniorum 62070 21
  • Mulleria tuzuniorum 62069 22
  • Mulleria tuzuniorum 62066 23
  • Mulleria tuzuniorum 62062 24
  • Mulleria tuzuniorum 62060 25
  • Mulleria tuzuniorum 62075 26
  • Mulleria tuzuniorum 62072 27
  • Mulleria tuzuniorum 62059 28
  • Mulleria tuzuniorum 62061 29
  • Mulleria tuzuniorum 62074 30
  • Mulleria tuzuniorum 62073 31
  • Mulleria tuzuniorum 62058 32
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount.

Set number: 2645

  • Mulleria tuzuniorum 62996 1
  • Mulleria tuzuniorum 62997 2
  • Mulleria tuzuniorum 62995 3
  • Mulleria tuzuniorum 62994 4
  • Mulleria tuzuniorum 62988 5
  • Mulleria tuzuniorum 62991 6
  • Mulleria tuzuniorum 62992 7
  • Mulleria tuzuniorum 62975 8
  • Mulleria tuzuniorum 62989 9
  • Mulleria tuzuniorum 62990 10
  • Mulleria tuzuniorum 62993 11
  • Mulleria tuzuniorum 62974 12
  • Mulleria tuzuniorum 63009 13
  • Mulleria tuzuniorum 63005 14
  • Mulleria tuzuniorum 62984 15
  • Mulleria tuzuniorum 62979 16
  • Mulleria tuzuniorum 63008 17
  • Mulleria tuzuniorum 63006 18
  • Mulleria tuzuniorum 62983 19
  • Mulleria tuzuniorum 62978 20
  • Mulleria tuzuniorum 63004 21
  • Mulleria tuzuniorum 63000 22
  • Mulleria tuzuniorum 62976 23
  • Mulleria tuzuniorum 62981 24
  • Mulleria tuzuniorum 63003 25
  • Mulleria tuzuniorum 62998 26
  • Mulleria tuzuniorum 62977 27
  • Mulleria tuzuniorum 62982 28
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount.

Set number: 2643

  • Mulleria tuzuniorum 53725 1
  • Mulleria tuzuniorum 53721 2
  • Mulleria tuzuniorum 53722 3
  • Mulleria tuzuniorum 53719 4
  • Mulleria tuzuniorum 53724 5
  • Mulleria tuzuniorum 53720 6
  • Mulleria tuzuniorum 53723 7
  • Mulleria tuzuniorum 53717 8
  • Mulleria tuzuniorum 53713 9
  • Mulleria tuzuniorum 53712 10
  • Mulleria tuzuniorum 53710 11
  • Mulleria tuzuniorum 53711 12
  • Mulleria tuzuniorum 53707 13
  • Mulleria tuzuniorum 53708 14
    10µm
Carlamuelleria tuezueniorum, Middle Paleocene, DSDP Leg 39, Sites 354, Ceará Rise, North Atlantic Ocean

In the mobile mount.

Set number: 2641

  • Mulleria tuzuniorum 68104 1
  • Mulleria tuzuniorum 68103 2
  • Mulleria tuzuniorum 68077 3
  • Mulleria tuzuniorum 68074 4
  • Mulleria tuzuniorum 68105 5
  • Mulleria tuzuniorum 68101 6
  • Mulleria tuzuniorum 68076 7
  • Mulleria tuzuniorum 68075 8
  • Mulleria tuzuniorum 68073 9
  • Mulleria tuzuniorum 68072 10
  • Mulleria tuzuniorum 68071 11
  • Mulleria tuzuniorum 68069 12
  • Mulleria tuzuniorum 68097 13
  • Mulleria tuzuniorum 68093 14
  • Mulleria tuzuniorum 68098 15
  • Mulleria tuzuniorum 68096 16
  • Mulleria tuzuniorum 68100 17
  • Mulleria tuzuniorum 68094 18
  • Mulleria tuzuniorum 68099 19
  • Mulleria tuzuniorum 68095 20
  • Mulleria tuzuniorum 68092 21
  • Mulleria tuzuniorum 68091 22
  • Mulleria tuzuniorum 68089 23
  • Mulleria tuzuniorum 68090 24
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, IODP Leg 171, Site 1051A, Black Nose, Western North Atlantic

In the mobile mount.

Set number: 2640

  • Mulleria tuzuniorum 54488 1
  • Mulleria tuzuniorum 54490 2
  • Mulleria tuzuniorum 54477 3
  • Mulleria tuzuniorum 54485 4
  • Mulleria tuzuniorum 54487 5
  • Mulleria tuzuniorum 54489 6
  • Mulleria tuzuniorum 54476 7
  • Mulleria tuzuniorum 54486 8
  • Mulleria tuzuniorum 54482 9
  • Mulleria tuzuniorum 54483 10
  • Mulleria tuzuniorum 54481 11
  • Mulleria tuzuniorum 54480 12
    10µm
Carlamuelleria tuezueniorum, Late Paleocene, DSDP Leg 39, Sites 354, Ceará Rise, North Atlantic Ocean

In the mobile mount.

Final Epithet
Carlamuelleria tuezueniorum Varol, 2023
Basionym

Carlamuelleria tuezueniorum Varol, 2023

Diagnosis

Large (8.0–12.0 μm) species of Carlamuelleria possessing a non-birefringent distal shield with 23 to 40 segments, a birefringent tube cycle with the often serrated periphery and a vestigial, diminutive proximal shield.

Etymology

In honour of Dinçer Ates¸ Tüzün and Ruhiye Tüzün, Geologists, Ankara, Turkey.

Dimensions Of Holotype

Diameter of the distal shield: 9.43 μm (measured in plan view in phase-contrast), the maximum thickness of the distal shield: 3.97 μm (measured in side view in cross-polarised light), the diameter of the tube cycle: 3.88 μm (measured in plan view in phase-contrast).

Description

The distal shield is lenticular in shape, having a shallow depression on its less convex side, and consists of about 23–40 segments. The tube cycle and vestigial diminutive proximal shield combination resembles a brandy goblet-shaped structure in side view. The tube cycle is often flush with the distal end of the shield but extends above it by a fraction (Plate 7, Figs. 4, 8, and 10 in Varol, 2023).

Optical Properties

In cross-polarised light, identifying the curved direction of the extinction lines is difficult. The inclined (oblique) extinction lines of the proximal side. With the gypsum plate inserted, the blue interference colours are diagonally aligned on the proximal and distal sides of the tube cycle (P7. 4, Figs. 14, 16; Plate8, 14, 16 in Varol, 2023). The blue sectors of the tube cycle and shield are aligned in opposite directions. The entire specimen appears birefringent in side view.

Remarks

In plan view, Carlamuelleria tuezueniorum is distinguished from Carlamuelleria bramlettei by having more segments in its shield (>23 segments). It differs from the subcircular Carlamuelleria karacuhae by having a circular tube cycle. In contrast, the latter has an elliptical tube cycle.

Carlamuelleria tuezueniorum is easily identified from Davidbukrya species in side view by having a brandy goblet-shaped structure formed by the arrangement of the tube cycle and the vestigial, diminutive proximal shield. The proximal shield is absent in Davidbukrya.

After mapping its optical properties, Carlamuelleria tuezueniorum is differentiated from Davidbukrya and Markalius in plan view by having a diagonal interference colour alignment in the tube cycle. In contrast, in their tube cycle, Davidbukrya and Markalius exhibit an axial interference colour alignment (i.e., a blue sector horizontal on the distal side and vertical on the proximal side). Moreover, Carlamuelleria tuezueniorum is consistently and confidently differentiated from Markalius in side view by having a vestigial, diminutive proximal shield, whereas the latter has a spaced proximal shield.

References

Varol, O. 2023. New paleocene calcareous nannofossils: Carlamuelleria, Davidbukrya, Mauriceblackia, Noelia and Senelia. Marine Micropaleontology. 180: 1-41.