Set number: 2446

  • Heliolithus margarita 61170 1
  • Heliolithus margarita 61166 2
  • Heliolithus margarita 61153 3
  • Heliolithus margarita 61142 4
  • Heliolithus margarita 61171 5
  • Heliolithus margarita 61167 6
  • Heliolithus margarita 61152 7
  • Heliolithus margarita 61141 8
  • Heliolithus margarita 61149 9
  • Heliolithus margarita 61150 10
  • Heliolithus margarita 61169 11
  • Heliolithus margarita 61147 12
  • Heliolithus margarita 61151 13
  • Heliolithus margarita 61148 14
  • Heliolithus margarita 61168 15
  • Heliolithus margarita 61145 16
  • Heliolithus margarita 61143 17
  • Heliolithus margarita 61165 18
  • Heliolithus margarita 61174 19
  • Heliolithus margarita 61157 20
  • Heliolithus margarita 61144 21
  • Heliolithus margarita 61164 22
  • Heliolithus margarita 61172 23
  • Heliolithus margarita 61156 24
  • Heliolithus margarita 61136 25
  • Heliolithus margarita 61159 26
  • Heliolithus margarita 61162 27
  • Heliolithus margarita 61161 28
  • Heliolithus margarita 61137 29
  • Heliolithus margarita 61158 30
  • Heliolithus margarita 61163 31
  • Heliolithus margarita 61160 32
  • Heliolithus margarita 61140 33
  • Heliolithus margarita 61138 34
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Holotype in the mobile mount

Set number: 2445

  • Heliolithus margarita 61238 1
  • Heliolithus margarita 61237 2
  • Heliolithus margarita 61234 3
  • Heliolithus margarita 61233 4
  • Heliolithus margarita 61239 5
  • Heliolithus margarita 61236 6
  • Heliolithus margarita 61235 7
  • Heliolithus margarita 61232 8
  • Heliolithus margarita 61225 9
  • Heliolithus margarita 61222 10
  • Heliolithus margarita 61229 11
  • Heliolithus margarita 61226 12
  • Heliolithus margarita 61224 13
  • Heliolithus margarita 61223 14
  • Heliolithus margarita 61228 15
  • Heliolithus margarita 61227 16
  • Heliolithus margarita 61230 17
  • Heliolithus margarita 61231 18
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount.

Set number: 2444

  • Heliolithus margarita 61769 1
  • Heliolithus margarita 61766 2
  • Heliolithus margarita 61765 3
  • Heliolithus margarita 61751 4
  • Heliolithus margarita 61770 5
  • Heliolithus margarita 61767 6
  • Heliolithus margarita 61764 7
  • Heliolithus margarita 61752 8
  • Heliolithus margarita 61757 9
  • Heliolithus margarita 61756 10
  • Heliolithus margarita 61762 11
  • Heliolithus margarita 61761 12
  • Heliolithus margarita 61759 13
  • Heliolithus margarita 61758 14
  • Heliolithus margarita 61763 15
  • Heliolithus margarita 61760 16
  • Heliolithus margarita 61755 17
  • Heliolithus margarita 61753 18
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2443

  • Heliolithus margarita 62158 1
  • Heliolithus margarita 62162 2
  • Heliolithus margarita 62166 3
  • Heliolithus margarita 62161 4
  • Heliolithus margarita 62159 5
  • Heliolithus margarita 62163 6
  • Heliolithus margarita 62160 7
  • Heliolithus margarita 62164 8
  • Heliolithus margarita 62175 9
  • Heliolithus margarita 62170 10
  • Heliolithus margarita 62171 11
  • Heliolithus margarita 62167 12
  • Heliolithus margarita 62176 13
  • Heliolithus margarita 62169 14
  • Heliolithus margarita 62172 15
  • Heliolithus margarita 62168 16
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2442

  • Heliolithus margarita 62473 1
  • Heliolithus margarita 62472 2
  • Heliolithus margarita 62457 3
  • Heliolithus margarita 62456 4
  • Heliolithus margarita 62498 5
  • Heliolithus margarita 62497 6
  • Heliolithus margarita 62495 7
  • Heliolithus margarita 62493 8
  • Heliolithus margarita 62491 9
  • Heliolithus margarita 62490 10
  • Heliolithus margarita 62462 11
  • Heliolithus margarita 62458 12
  • Heliolithus margarita 62492 13
  • Heliolithus margarita 62489 14
  • Heliolithus margarita 62461 15
  • Heliolithus margarita 62460 16
  • Heliolithus margarita 62487 17
  • Heliolithus margarita 62488 18
  • Heliolithus margarita 62474 19
  • Heliolithus margarita 62475 20
  • Heliolithus margarita 62483 21
  • Heliolithus margarita 62482 22
  • Heliolithus margarita 62471 23
  • Heliolithus margarita 62468 24
  • Heliolithus margarita 62485 25
  • Heliolithus margarita 62484 26
  • Heliolithus margarita 62470 27
  • Heliolithus margarita 62469 28
  • Heliolithus margarita 62477 29
  • Heliolithus margarita 62467 30
  • Heliolithus margarita 62466 31
  • Heliolithus margarita 62486 32
  • Heliolithus margarita 62479 33
  • Heliolithus margarita 62465 34
  • Heliolithus margarita 62464 35
  • Heliolithus margarita 62481 36
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount

Set number: 2441

  • Heliolithus margarita 64807 1
  • Heliolithus margarita 64799 2
  • Heliolithus margarita 64800 3
  • Heliolithus margarita 64803 4
  • Heliolithus margarita 64806 5
  • Heliolithus margarita 64798 6
  • Heliolithus margarita 64801 7
  • Heliolithus margarita 64802 8
  • Heliolithus margarita 64796 9
  • Heliolithus margarita 64804 10
  • Heliolithus margarita 64813 11
  • Heliolithus margarita 64810 12
  • Heliolithus margarita 64797 13
  • Heliolithus margarita 64805 14
  • Heliolithus margarita 64812 15
  • Heliolithus margarita 64811 16
  • Heliolithus margarita 64808 17
  • Heliolithus margarita 64809 18
    10µm
Heliolithus margarita, Middle Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2440

  • Heliolithus margarita 66016 1
  • Heliolithus margarita 66025 2
  • Heliolithus margarita 66013 3
  • Heliolithus margarita 66028 4
  • Heliolithus margarita 66015 5
  • Heliolithus margarita 66026 6
  • Heliolithus margarita 66014 7
  • Heliolithus margarita 66027 8
  • Heliolithus margarita 66024 9
  • Heliolithus margarita 66020 10
  • Heliolithus margarita 66032 11
  • Heliolithus margarita 66029 12
  • Heliolithus margarita 66022 13
  • Heliolithus margarita 66019 14
  • Heliolithus margarita 66031 15
  • Heliolithus margarita 66030 16
  • Heliolithus margarita 66034 17
  • Heliolithus margarita 66033 18
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount

Set number: 2439

  • Heliolithus margarita 66737 1
  • Heliolithus margarita 66735 2
  • Heliolithus margarita 66729 3
  • Heliolithus margarita 66732 4
  • Heliolithus margarita 66736 5
  • Heliolithus margarita 66734 6
  • Heliolithus margarita 66730 7
  • Heliolithus margarita 66731 8
  • Heliolithus margarita 66721 9
  • Heliolithus margarita 66728 10
  • Heliolithus margarita 66725 11
  • Heliolithus margarita 66724 12
  • Heliolithus margarita 66722 13
  • Heliolithus margarita 66727 14
  • Heliolithus margarita 66726 15
  • Heliolithus margarita 66723 16
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

In the mobile mount

Set number: 2438

  • Heliolithus margarita 69390 1
  • Heliolithus margarita 69391 2
  • Heliolithus margarita 69381 3
  • Heliolithus margarita 69394 4
  • Heliolithus margarita 69392 5
  • Heliolithus margarita 69389 6
  • Heliolithus margarita 69382 7
  • Heliolithus margarita 69395 8
  • Heliolithus margarita 69388 9
  • Heliolithus margarita 69386 10
  • Heliolithus margarita 69387 11
  • Heliolithus margarita 69385 12
  • Heliolithus margarita 69383 13
    10µm
Heliolithus margarita, Late Paleocene, IODP Leg 171, Site 1051A, Black Nose, Western North Atlantic

In the mobile mount

Set number: 2477

  • Heliolithus margarita 22385 1
  • Heliolithus margarita 22384 2
  • Heliolithus margarita 22386 3
  • Heliolithus margarita 22387 4
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2476

  • Heliolithus margarita 22060 1
  • Heliolithus margarita 22061 2
  • Heliolithus margarita 22062 3
  • Heliolithus margarita 22063 4
    10µm
Heliolithus margarita, Middle Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2475

  • Heliolithus margarita 21107 1
  • Heliolithus margarita 21106 2
  • Heliolithus margarita 21109 3
  • Heliolithus margarita 21108 4
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2474

  • Heliolithus margarita 20972 1
  • Heliolithus margarita 20971 2
  • Heliolithus margarita 20974 3
  • Heliolithus margarita 20973 4
  • Heliolithus margarita 20976 5
  • Heliolithus margarita 20975 6
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean

Set number: 2473

  • Heliolithus margarita 20899 1
  • Heliolithus margarita 20900 2
  • Heliolithus margarita 20902 3
  • Heliolithus margarita 20901 4
    10µm
Heliolithus margarita, Late Paleocene, DSDP Leg 25, Site 245, Mozambique Channel (Davie Ridge), Indian Ocean
Final Epithet
Heliolithus margarita Bowman & Varol, 2021
Basionym

Heliolithus? margarita Bowman & Varol, 2021

Diagnosis

A medium to large-sized and circular species of Heliolithus featuring a strangely formed disc and a vestigial and/or undetectable column. The relatively wide central canal is closed by the plug/tube cycle. In plan view, the bizarre disc gives a “daisy-like” appearance. The species appears birefringent in all views.

Etymology

From Latin margarita, meaning daisy—referring to the daisy-like appearance of the species in plan-view.

Dimensions Of Holotype

Overall height: 3.61μm; Disc diameter: 9.56μm.

Description

Medium to large (6.0–10.0μm) circular species composed of one disc and a vestigial column [very low or undetectable column]. The disc is moderately wide and appears relatively flat or very slightly convex. The relatively wide central canal is closed by a plug/tube cycle but often removed by adverse paleoenvironmental conditions. The disc is comprised of about 24–40 segments, but the segments of the plug/tube cycle are indistinct.

Optical Properties

A distinct depression is present on one side of the disc. In plan view, the disc's character produces a daisy-like appearance in the species. The distal side is associated with dextrogyre extinction lines (e.g. Pl. 79, figs. 10, 14 in Bowman & Varol, 2021), whereas laevogyre extinction lines characterise the proximal side (e.g. Pl. 79, figs. 11–15 in Bowman & Varol, 2021). The horizontal axis relates to the blue sector on the distal side; however, the vertical axis relates to the blue sector on the proximal side (observed using the gypsum plate). The disc and column are birefringent in side-view and plan-view. More specifically, the disc and column show a whiteish-yellowish birefringence colour in plan-view, and the disc displays a reddish-yellowish birefringence colour in the side-view.

Remarks

Heliolithus? margarita represents a strange and atypical form; therefore, assigning this species to Heliolithus is questionable. Heliolithus? margarita possesses a vestigial column, which is the diagnostic feature used to separate this species from other Heliolithus species. In addition, this species differs from the species of Caycedoa based on the diagnostic birefringent pattern of its disc. The disc is non-birefringent in plan-view in the species of Caycedoa.

References

Bowman, A. R. & Varol, O. 2021. A Taxonomic Revision of Heliolithaceae - Applications in Resolving the Problematic Calcareous Nannofossil Biostratigraphy of the Paleocene. In: M. Montenary, M. (Ed.). Calcareous nannofossil biostratigraphy of the Stratigraphy and timescales. 6: 43-223.