Type Species

Ceratolithus amplificus Bukry and Percival, 1971


Non-birefringent Ceratolith having a C-axis is perpendicular to the plane of a horseshoe or natural settling of the coccolith. Having ornamented straight horn and a non-ornamented curved horn. The apical area is well developed.


Nicklithus differs from Ceratolithus Kamptner (1950) by being non-birefringent, whereas the latter is strongly birefringent. It is distinguished from Amaurolithus Gartner & Bukry (1975) by having a straight horn with keels.


Bukry, D. & Percival, S. F., (1971): New Tertiary calcareous nannofossils. -Tulane Studies in Geology and Paleontology. 8: 123-146.

Gartner, S. & Bukry, D., (1975): Morphology and phylogeny of the coccolithophycean family Ceratolithaceae. - Journal of Research of the U.S. Geological Survey. 3: 451-465.

Kamptner, E., (1950): Über den submikroskopischen Aufbau der Coccolithen. - Anzeiger der Österreichische Akademie der Wissenschaften, Mathematische-Naturwissenschafliche Klasse, Wien. 87: 152-158.

Raffi, I., Backman, J. & Rio, D., (1998): Evolutionary trends of tropical calcareous nannofossils in the late Neogene. - Marine Micropaleontology. 35(1): 17-41