Pennalithus rugosus

(Bramlette & Wilcoxon, 1967) Varol, 2025 May 17, 2021
P rugosus 1

Set number: 1424

  • Pennalithus rugosus VR 10055 1
  • Pennalithus rugosus VR 10056 2
  • Pennalithus rugosus VR 10059 3
  • Pennalithus rugosus VR 10048 4
  • Pennalithus rugosus VR 10058 5
  • Pennalithus rugosus VR 10057 6
  • Pennalithus rugosus VR 10053 7
  • Pennalithus rugosus VR 10035 8
  • Pennalithus rugosus VR 10052 9
  • Pennalithus rugosus VR 10047 10
  • Pennalithus rugosus VR 10050 11
  • Pennalithus rugosus VR 10045 12
  • Pennalithus rugosus VR 10039 13
  • Pennalithus rugosus VR 10038 14
  • Pennalithus rugosus VR 10037 15
  • Pennalithus rugosus VR 10041 16
    10µm
Pennalithus rugosus, Late Miocene, DSDP Leg 23, Site 225, Red Sea

Set number: 3814

  • Pennalithus rugosus VR 16456 1
  • Pennalithus rugosus VR 16455 2
  • Pennalithus rugosus VR 16453 3
  • Pennalithus rugosus VR 16458 4
  • Pennalithus rugosus VR 16457 5
  • Pennalithus rugosus VR 16454 6
  • Pennalithus rugosus VR 16452 7
  • Pennalithus rugosus VR 16459 8
  • Pennalithus rugosus VR 16446 9
  • Pennalithus rugosus VR 16445 10
  • Pennalithus rugosus VR 16447 11
  • Pennalithus rugosus VR 16461 12
  • Pennalithus rugosus VR 16451 13
  • Pennalithus rugosus VR 16450 14
  • Pennalithus rugosus VR 16448 15
  • Pennalithus rugosus VR 16460 16
    10µm
Pennalithus rugosus, Late Miocene, ODP Leg 154, Hole 925A, Ceará Rise, Equatorial Atlantic Ocean

Set number: 3815

  • Pennalithus rugosus VR 10389 1
  • Pennalithus rugosus VR 10393 2
  • Pennalithus rugosus VR 10395 3
  • Pennalithus rugosus VR 10400 4
  • Pennalithus rugosus VR 10390 5
  • Pennalithus rugosus VR 10392 6
  • Pennalithus rugosus VR 10396 7
  • Pennalithus rugosus VR 10399 8
  • Pennalithus rugosus VR 10388 9
  • Pennalithus rugosus VR 10387 10
  • Pennalithus rugosus VR 10385 11
  • Pennalithus rugosus VR 10398 12
    10µm
Pennalithus rugosus, Early Pliocene, DSDP Leg 23, Site 225, Red Sea

Set number: 3812

  • Pennalithus rugosus VR 19060 1
  • Pennalithus rugosus VR 19076 2
  • Pennalithus rugosus VR 19056 3
  • Pennalithus rugosus VR 19077 4
  • Pennalithus rugosus VR 19058 5
  • Pennalithus rugosus VR 19054 6
  • Pennalithus rugosus VR 19055 7
  • Pennalithus rugosus VR 19053 8
  • Pennalithus rugosus VR 19069 9
  • Pennalithus rugosus VR 19064 10
  • Pennalithus rugosus VR 19061 11
  • Pennalithus rugosus VR 19070 12
  • Pennalithus rugosus VR 19068 13
  • Pennalithus rugosus VR 19063 14
  • Pennalithus rugosus VR 19062 15
  • Pennalithus rugosus VR 19071 16
  • Pennalithus rugosus VR 19052 17
  • Pennalithus rugosus VR 19074 18
  • Pennalithus rugosus VR 19075 19
  • Pennalithus rugosus VR 19078 20
  • Pennalithus rugosus VR 19065 21
  • Pennalithus rugosus VR 19073 22
  • Pennalithus rugosus VR 19057 23
  • Pennalithus rugosus VR 19087 24
  • Pennalithus rugosus VR 19067 25
  • Pennalithus rugosus VR 19072 26
  • Pennalithus rugosus VR 19059 27
  • Pennalithus rugosus VR 19086 28
    10µm
Pennalithus rugosus, Late Miocene, ODP Leg 154, Hole 926A, Ceará Rise, western equatorial Atlantic.

Set number: 3813

  • Pennalithus rugosus VR 18784 1
  • Pennalithus rugosus VR 18785 2
  • Pennalithus rugosus VR 18788 3
  • Pennalithus rugosus VR 18765 4
  • Pennalithus rugosus VR 18783 5
  • Pennalithus rugosus VR 18786 6
  • Pennalithus rugosus VR 18787 7
  • Pennalithus rugosus VR 18789 8
  • Pennalithus rugosus VR 18770 9
  • Pennalithus rugosus VR 18771 10
  • Pennalithus rugosus VR 18774 11
  • Pennalithus rugosus VR 18763 12
  • Pennalithus rugosus VR 18769 13
  • Pennalithus rugosus VR 18772 14
  • Pennalithus rugosus VR 18773 15
  • Pennalithus rugosus VR 18768 16
  • Pennalithus rugosus VR 18747 17
  • Pennalithus rugosus VR 18749 18
  • Pennalithus rugosus VR 18752 19
  • Pennalithus rugosus VR 18758 20
  • Pennalithus rugosus VR 18746 21
  • Pennalithus rugosus VR 18748 22
  • Pennalithus rugosus VR 18753 23
  • Pennalithus rugosus VR 18757 24
  • Pennalithus rugosus VR 18778 25
  • Pennalithus rugosus VR 18781 26
  • Pennalithus rugosus VR 18779 27
  • Pennalithus rugosus VR 18760 28
  • Pennalithus rugosus VR 18777 29
  • Pennalithus rugosus VR 18782 30
  • Pennalithus rugosus VR 18780 31
  • Pennalithus rugosus VR 18754 32
  • Pennalithus rugosus VR 18793 33
  • Pennalithus rugosus VR 18764 34
  • Pennalithus rugosus VR 18794 35
  • Pennalithus rugosus VR 18795 36
  • Pennalithus rugosus VR 18762 37
  • Pennalithus rugosus VR 18761 38
  • Pennalithus rugosus VR 18759 39
  • Pennalithus rugosus VR 18790 40
  • Pennalithus rugosus VR 18767 41
  • Pennalithus rugosus VR 18766 42
  • Pennalithus rugosus VR 18756 43
  • Pennalithus rugosus VR 18755 44
  • Pennalithus rugosus VR 18792 45
  • Pennalithus rugosus VR 18791 46
    10µm
Pennalithus rugosus, Late Miocene, ODP Leg 154, Hole 926A, Ceará Rise, western equatorial Atlantic.

Set number: 1019

  • Pennalithus rugosus 29760 1
  • Pennalithus rugosus 29759 2
  • Pennalithus rugosus 29756 3
  • Pennalithus rugosus 29757 4
    10µm
Pennalithus rugosus, Middle Miocene, Malta

Set number: 1018

  • Pennalithus rugosus 29766 1
  • Pennalithus rugosus 29761 2
  • Pennalithus rugosus 29763 3
  • Pennalithus rugosus 29768 4
  • Pennalithus rugosus 29765 5
  • Pennalithus rugosus 29762 6
  • Pennalithus rugosus 29764 7
  • Pennalithus rugosus 29767 8
    10µm
Pennalithus rugosus, Middle Miocene, Malta
Final Epithet
Pennalithus rugosus (Bramlette & Wilcoxon, 1967) Varol, 2025
Basionym

Triquetrorhabdulus rugosus Bramlette & Wilcoxon, 1967

Synonyms

Orthorhabdus rugosus (Bramlette & Wilcoxon, 1967) Young & Bown 2014

Description

Broad species of Pennalithus have two blades of unequal size in the same plane and a median ridge that is straight or gently curved.

Optical Properties: Pennalithus rugosus naturally settles in its plan view and appears in constant extinction (the optic axis is perpendicular relative to the microscope stage, Plate 19, Figs. 1-8, Plate 20, Figs. 1-8 in Varol, 2025). The natural settlement of its side view is sporadic; therefore, the optical properties of the side view are obtained from a mobile mount (Plate, 19 Figs. 9-24, Plate 20, Figs. 9-18 in Varol, 2025). It exhibits parallel extinction (extinction angle is 0°) and displays length-slow (+) elongation in the side view profile (Plate 19, Figs. 10, 14; Plate 20, Figs. 10,14, 17-18 in Varol, 2025).

Remarks

Pennalithus rugosus differs from Pennalithus rioi by being broader and having two blades distinctly different in width compared to Pennalithus rioi.

References

Bramlette, M. N. & Wilcoxon, J. A. 1967. Middle Tertiary calcareous nannoplankton of the Cipero section, Trinidad, W.I. Tulane Studies in Geology and Paleontology 5: 93-131.

Varol, O. 2025a. A practical guide to optical studies of calcareous nannofossils. Grzybowski Foundation Special Publication. 29: 1-222

Young, J. R. & Bown, P. R. 2014. Some emendments to calcareous nannoplankton taxonomy. Journal of Nannoplankton Research 33(1): 39-46.

Olafsson, G. 1989. Quantitative calcareous nannofossil biostratigraphy of upper Oligocene to middle Miocene sediment from ODP Hole 667A and Middle Miocene sediment from DSDP Site 574. Proceedings of the Ocean Drilling Program, Scientific Results 108: 9-22.

Taxonomy
Pennalithus
Biostratigraphy
  • Phanerozoic
    • Cenozoic
      • Neogene
        • Pliocene
          • Early Pliocene
            • Zanclean
        • Miocene
          • Middle Miocene
            • Serravalian
Stratigraphic Range
Serravalian - Zanclean
Age (Ma)
13.33 - 5.23
Holotype
Pennalithus rugosus 1075 16

Triquetrorhabdulus rugosus

Late Miocene

Cipero Section, Trinidad